It appears Sony's wholly owned studio Naughty Dog, famous for the Jak & Daxter, Uncharted, and The Last of Us franchises, has lost yet another lead developer.

The Last of Us lead character artist Michael Knowland updated his LinkedIn page (spotted by Superannuation) to say his tenure at Naughty Dog ended, or will end, sometime in April 2014. It's unclear if Knowland has actually left Naughty Dog, and we've reached out to a representative for the company for more information.

This news comes after recent departures of a handful of high-ranking Naughty Dog employees, including Uncharted 4 director Justin Richmond, writer Amy Hennig, lead artist Nate Wells, and actor Todd Stashwick.

Of course, it's not out of the ordinary for developers to leave their job for a new opportunity, but the sheer number of employees who have left Naughty Dog in recent days is certainly something to take note of. For more on the changes at Naughty Dog and other first-party Sony studios, be sure to read GameSpot's editorial on what the turnover might mean.